Full text - Acta Palaeontologica Polonica

Full text - Acta Palaeontologica Polonica

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No. 2
HALSZKA OSMOLSKA
FAMENNIAN PHACOPIDAE FROM THE HOLY CROSS MOUNTAINS
(POLAND)
Abstract. - Nine species of the Famennian phacopid trilobites are described from
the southern part of the Holy Cross Mts. In addition to forms already recorded
from this area the writer describes some species never before found in Poland, as
well as two quite new species, namely: Trimerocephalus polonicus and Dianops?
trifolius.
INTRODUCTION
The described material was collected in 1948 and 1949 by Dr. Zofia
Kielan, from Kadzielnia, Psiarnia, Herby (within the precincts of town
Kielce), Lagow, Galezice, Kowala and Jablonna near Bork6w, all lying
in the southern part of the Holy Cross Mountains.
The terminology used in descriptions of glabella is partly that
proposed by V. Jaanusson (1956), to say: lobes and lateral furrows in
glabella have been lettered L (lobus) and S (sulcus) respectively, and
numbered postero-anteriorly. The term "preoccipital ring" has been used
to designate the posterior glabellar lobe (Lj) and that of "preoccipital
furrow" - to designate lateral glabellar furrow (SI)' "Preoccipital ring"
is a definition more appropriate with the morphological character of this
glabellar element in phacopids than that of "lobe", since in most cases,
similarly to the occipital ring, it is in the form of a uniform segment.
R. & E. Richter (1926) introduced the following terms in their descriptions of phacopid trilobites: "Zwischenring" = "basal lobe" = "preoccipital ring" = L, and "Zwischenfurche" = "preoccipital furrow" = SI'
Instead of Delo's (1940, s. 15) term "subcranial furrow", the more correct
term "subcranidial furrow" is used as an equivalent to R. & E. Richter
.IVerschlussfurche".
The abbreviations used here are: tr. - transversal i. e. perpendicular
to the axis of symmetry; sag. - sagittal i. e. parallel to the longitudinal
axis. Length of glabella has been measured without the occipital ring.
Famennian phacopids have been described from the Holy Cross
Mountains by several authors, i. e. by C. F. Roemer (1866), G. Giirich
120
HALSZKA OSM6LSKA
(1896), D. Sobolev (1911), R. & E. Richter (1926) and also mentioned in
papers by the Polish geologists: J . Czarnocki (1948 and elsewhere) and
J . Samsonowicz (1917 and elsewhere) .
C. F . Ro emer (1866) has described speci es Dianops typhlops (Giirich ,
189fl) under the n ame of Phacops cry pto pht alm us (Ernmrich).
G . Giirich (1896) gave this species the new n ame of Trimerocephalus
t ?lphlops. In t he sa me pape r h e al so d escribed other Famennian phacopids
suc h as Ph acop s posid oniae n. sp . = Phacops granulatus (Munster, 1840) ,
Triuierocephalus sp. (in cisu s? A . Roemer , 1866) = N ephranops incisus
'in cisus (A . Roemer, 1866) , Ph acops caecus n. sp. = T rimerocephalus cae cu s
(G tirich , 1896).
D . Sobol ev (1911) repor ts fro m Famennian beds of the Holy Cross
Mt s. the occurrence of Trimero cephalus caecus (Gurich , 1896) under the
name of P hacops su lcatus? Dr ev ermann , 1901.
R. & E. Rich ter (1926), in their work o n Upper D evonian t r ilobites,
amo ng ot he rs , also desc ribe Famen nian p hacopids occur r in g in Poland
an d elsew here .
To the li st of -ph acopid s h it herto reco rde d f rom P oland t he pr esent
writer adds two n ew species: Trimerocephalus polonicu s n . s p. and Dia nop s? tri folius n . sp ., as w ell as som e spe cies known from Great Britain ,
Germa ny a nd U. S. S. R. , but n ever u p to now found in Poland, to say :
Phacops w edekin di wedekindi R. & E. Richter, 1926 ; Trimerocephalus
ma stoph talmus (Reinh. Richter , 1856) ; Ductina ductifrons (R. & E. Richter, 1923) . Specimen s of N ephranop s incisu s incisus (A . Roem., 1866) were
not available to t he writer. On e damaged cephalon of this form, found
by Giirich in Kielce , is referabl e to lowest F amennian deposits and perha ps eve n to t hose of the upp ermost Frasnian (Mant icocera s zone) , as
surmised by R. & E. Richt er.
All the h ere de scr ibed spe cimens a re d epo sited in the Geological
S urvey Museum in Warsaw (Instytut Geologiczny) , referred to by the abbr eviated form I . G .
L aboratory work du ring the preparation of t his pa pe r has been
ca r ried out by the writer in 1954 , a t the Institute of Paleozoology of the
Polish Academy of Sciences , under th e guidance of Prof. Dr. R. Kozlowski whose valuable suggestions w ere of great assistance. Thanks are also
due to the following persons: to Dr. Z. Kielan for the privilege of d escribing her collection , for helping the author and for reading her
manuscript ; to Mrs. M . Pajchel for general information on th e geology of
the studied area; to Mrs. J. Humnicka for the English translation of the
present- paper; to Miss M. Czarnocka for th e pains taken in doing the photographs published .in:J hJs , paper.
FAMENNIAN PHACOPIDAE F R O M HOLY C R O SS MTS .
121
R EMA R K S ON THE GE OLOGY O F FAMENNIAN BEDS IN THE S OUTHERN P A nT
OF THE HOLY CROSS MTS.
Th e F amennian ph acopids d escribed in the present paper h av e been
coll ected from the following localities: Kadzielnia , P siarnia, H erby , Lagow ,
Ga le zice, Kowala and Jablonna (see fig. 1). All t hese site s are sit uated
so ut h of the Holy Cross Mts. r ange an d m ay b e grouped into t wo regio ns
of differen t lithologi cal an d fa unal cha ra cter: I. Cen tral (or K ielceLagow) synclinorium, II. Gal ezice-Dal eszyce synclin e.
Within th e central syncli no rium t he Famennian is represen ted by
a se ries of grey-yellowish s hale limestone d eposit s, ove r 100 m in th ickness. Th e trilobite fa una here is fa irly abunda n t, con sisting m ainly of
bli nd phacopid fo rms (se e tabl e of di st ributi on on p. 124) , suc h as Trim erocephalus caecus (Giirich) , T rim . pol oni cus n. sp., Trim . mastophtalm us
(Reinh . Richter) , Dianops t y phlops (Giirich) , Dianop s? trifo lius n. sp.,
Ductina du ctifrons R. & E . Richt er. Proeti d t r ilo bites are very rare.
Depos its of this typ e occ ur in the following localities:
Kadzi elnia In Kadzielnia Famen nia n beds r est on Frasnian
lim esto nes a nd ar e made up of marly shales inter bed d ed by limeston es .
This is a t hi ck s eries of depo sits containing a trilobite and lamellibra nch
fa un a, with f ragm en tary r emains of clymenid s in a poo r sta te of preser vat ion . After Czarnocki (1948) Cheilocer as and Clymenia zon es occur here .
Th E. foll owing ph acopids h av e been record ed: Trimero cephalus caecus
(Giir ich), Trim . mastophtalmus (Reinh. Richt er), Tr im. polonicus n . sp.,
Dianops? trifolius n . sp .
Psiarnia - The Famennian is dev eloped as marly sha les intercal ated
by limestones. Phacopid t r ilob ites enco un ter ed here are: Phacops granulatus (Munster) 1 , Dianops typhlops (Giirich) , Ductina du ctifrons (R. &
E. Richter) .
Herby - Black limestones outcrop on the roadway leading to Czarnow, containing Dianops typhlops (Giirich) .
Lagow - The Famennian outcrops occur within it s northernmost
a r ea , on th e va ll ey slope s of the Lagowica stream. Th ey are strongly
bituminous shale limestone deposits. The dark-grey Clymenia limestones
here occasionally yield a profusion of exoskeletons belonging to Trimeroceph alus caecus (Giirich) (see pl . II , fig. 3), those of Trim. mastophtalmus
(Reinh. Richter) being very scarce. Dianops typhlops (Gurich) is another
phacopid form here encountered .
1 The occurrence in Psiarnia of Phacops grannlatus (MUnst er) seems doubtful.
Its only record from this locality is that cited by Giirich (1896), but it has never
after been found here.
....
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Distribution of the Famennian in the southern part of the Holy Cross Mountains.
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FAMENNIAN PHACOPIDAE FROM HOLY CROSS MTS .
123
The facial development of the Famennian beds in Lagow slightly
d iffe rs from the type shale limestone facies on account of the predomina n ce of limestones over shales. The fa unal assemblage here is different
too. In addition to phacopids, very cop ious here, trilobites of the Proetidae
family are also represented.
Within t he Galezice-Daleszyce syncl ine Famennian beds are of
a limest on e faci es, frequently red coloured and characterized by sm all
thickness, never exceeding 30 m. The fa un a here is v ery rich and in a n
ex cellen t state of pres ervation. Of phacop id trilobites only two s pe cies
occur: Pha cops granulatus (Munster) and Ph. wedekindi w edekin di R. &
E. Richter, both pro vid ed with eyes, while proetids, nearly altoget h er
a bsen t from the central synclinorium, are h ere very numerous. Famenni an d ep osits of the limestone type occu r in the follow in g localities:
Gal ezice - Beds of F r asnian age absen t . The F amennian deposits
rest directly on A m phipora Givetian limest ones a nd consist of hard limestones displaying black, grey, pink and green ish coloura tion. After Czarnocki (1928) the Ch eil ocera s horizon is mis sing here, w hile the Upper
Famennian contains all the Clymen ia horizon s, those of Wocklumeria a nd
Gatte ndor fi a incl uded. The fauna is a markedly r ich one, w ith goniatites ,
cly rnenids, orthoceratids, brachiopods, lamellibranchs, trilo bites a nd fis h
r em ains. Of the phacopids we note h ere Pha cops granul atus (Munste r ).
Higher up are d eposits of Lower Carboniferous a ge (the Culm a nd Carbon iferous limestone lenses). Due to the taper ing of some of the beds, thickn es s of ou tcrops r anges fr om 3 to 10 m.
Ko w ala - The F am ennian b eds are r epresented h ere by nod ulous
limestones , often with a r eddish hue, overlyin g Frasnian limestones. After
Cza rnocki (1957) all the Clymenia horizons ar e here present. Of t he
ph acop id s there oc curred: Phacops wedek indi wedekindi R. & Richter,
an d Dianops t y phlops (Gii rich). Culm shales occur higher up.
Jablonna n ear Borkou: - The F amennian h as been here discov ered
by the l at e J. Czarnocki, aft er World War II . His fi eld wor k on the
geology of the district was carried on through 1949 , but the info rmat ion
thus obtained has never been published. Famennian beds overlay here the
Frasnian limestones . Similarly as in Kowala , a complete F amennian succession is exposed, beginning with Cheiloceras beds and probably including all the Clymenia horizons. The deposits consist of limestones with red
or grey colouration, while the beds interpreted by Czarnocki as of W ocklumeria horizon are of green colour. The fauna is very rich. Thickness
about 30 m . Of phacopid trilobites the only form occurring here is Phacops
granulatus (Munster) . Representatives of this species occurring within
lower Clymenia horizons consist of forms having cephalons of the sire
HALSZKA OSM6LSKA
124
of a few millimeters only, while the same species from t he green Wock- ·
lumeria limestones are with cephalons up to 18 mm in size. Culm shales.
overlie the Famennian deposits .
The geology of the Famennian beds within the Holy Cross Mts. has
not, thus far , been described in full detail. Hence the writer has experienced difficulties in est a bli shing th e horizons which hav e yielded the
st udied t r ilobit es. Th e only differenti ation which it ha s been possible to·
mak e is that b etween th e old er, i. e. Cheiloceras beds, and the young er,
i. e . Clymenia . b ed s. Most of the known species here de scribed are also
reco r d ed from the same bed s in other countries. The only exception is
Ductina duct ifrons R. & E . Ri chter. This spe cies has be en reported by
its authors (1926) fr om th e Mant icoceras and Cheiloceras horizons , i. e.
from Upper Frasnian an d lowest Famennian b eds, while in the Holy
Cross Mts. it has been yi eld ed by the Clymenia beds (Upper Famennian)
of Psiarnia .
Distribution of th e consider ed phacopid speci es in outcr ops of th e south ern part
of th e Holy Cross Mountains
~~~::~I
Species
1. Phacops granulatus (Miins ter)
2. Phacops wedekindi w edek i n di R. &
E. Richter
3. Trirnerocephalu s caecus (Giirich)
4. T r imeroeephalu» ma stophtalrn us
(Reinh, Richter]
5.
6.
7.
8.
9.
Trimerocephalus polonicus n. sp.
Dianops typhlops [Giirtch ]
Dianops? trifolius n. sp.
Ductina ductifron s R. & E. Richter
Phacops sp.
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*Th e bracketed + sign in dica t es t hat this particular species has been r ecorded
from th at site by other autho rs . but that the present w ri ter ha s not en cou ntered
it in that respective outcrop.
125
FAMEN N IAN P H ACO P I D AE FROM H O L Y CROSS MTS .
SYST EMA T IC DES CRIPTIONS
Family Phacopidae Hawl e & Corda , 1847 (emend . Delo , 1935)
Subfamily Phacopinae Reed, 1905
G enus Phacops Emmrich, 1839
Phacops granu latus (Munster 1840)
(fig. 2 ; pl. I , fig. 1)
1896. Phacop s p osidonia e GUrich ; G. GUri ch , Da s
Palaozoicum..., p . 363, p l. 15,
fig. 5 a -d .
1926. P h aco p s (P h aco p s) granulatus (MUnster); R. &
E. Richter , Die T r il ob iten ...,
p . 137, pl . 8, fi g. 34- 39 (w i t h previous sy non y m y ).
1955. Phacops (Pha cop s) g r an u l at us (M Un s te r ); Z . A. Maksimova, Trilobity.." p . 136,
pl. 10, fi g. 3-11.
Material. - 4 fragme nts and one complete cephalon with exosk eleton ,
1 pygidium with exoskeleton, 1 pygidi um without exoskeleton - all from
the grey limestone Clymenia beds of Galezice; 4 fragment s of cephalon
and 1 pygidium with exosk ele ton from the r ed limeston e Clymenia beds
of Jablonna; 2 damaged cep ha lons with exoskeleton from the light-green
limestone W ocklumeria beds of Jablonna.
Dim ension s of 5 specimens (in mm) -
Specimens
~~ I
Length of cephalon
Width of cephalon
Leng th of glabell a
Width of gl a b ell a
W id t h of p reoccipital ring L 1 (sag .)
L en gth of e y e
Di stance of e y e f rom posterior
furrow
Angl e of dorsal f ur rows
L ength of pygid ium
Width of pygidiu m
. Length of rh achis
Width of r hachi s
.', :\ l lI S\ ' lIl11 Ca ut!0I!II(' N lI I II I H' I ~ o r
I. G .
169.II .2
see t able be low.
I. G .
169.II.1
I. G .
169.II. 4
I. G.
169.II.3
5.0
8.0
4.0
4.8
0.6
1.8
7.0
11.2
5.2
7.0
0.5
2.9
12.7
18.5
11.0
13.0
16.0
19.0
2.0
7.8
0.8
72 0
0.9
75°
0.8
65°
t he Ge o logica l In stitut e (I. C. )
5.1
I:G.*
169.II .5
2.1
3.5
7.5
3.1
1.7
i ll
\\' RI SIH \.
De scription. - Ou tlin e of cephalon semicircular. In longitudinal
section occipital r ing as high as glabella; anterior portion of gl ab ell a
overhanging the border. In transvers e section glabella st rongly v ault ed ,
eye lobe nearly vertical to palpebral lob e. Greatest -width of glabella be-
126
HALSZKA OS M6LSKA
fore it s mid-length, at level of anterior end of the eye; length/width ratioof glabella 0.8; dorsal furrows a t first st raight , diverging at the level of
the anterior end of eye ; angle of dorsal furrows 65° to 75° ; three fourths
of cheek area occupied by ey es , their anterior margin directly touching.
20
,'"
3
Fig. 2. - Pha cop s grallulatus (Mu nster): Ia cephalon , la te r a l view; I b sa me
spe cim e n, a n te rior view (1. G. 169 II. 1); 2a pygid ium, latera l v ie w ; 2b sa m e
specimen, posterior v iew (1. G. 169. II. \23).
the glabella; the ratio of the ey e length to its distance from posterior
border f ur row strongly variable. In the 7 specimens th at have b een
measured it is: 2.2, 3.2, 3.6, 3.7, 4.8, 5.8 and 6.4 , i. e. an average of 4.2.
Lenses on eye lobe round, hardly contacting one anothe r . In an individual
7 rom long and 11.2 rom wide the lenses are arranged in 13 vertical rows,.
3 lenses of 0.4 rom in each row; there are also some irregularly arranged
lenses along the posterior eye end. In other cephalons the number of
vertical rows is the same with 4 lenses in each row , in an alternate ar -·
rangern ent. Total number of l enses ranges from 39 to 52. P ygidium s ho rt
and stout , the length/width ratio being 0.39.
Ornamentation of glabella consists of large, densely arranged tubercula, smaller above the anterior border furrow a nd in the cheek area ;
lateral border on the fixed cheeks with minute granulation; pygidiaL
rhachis covered by small and rather s pa r se tubercles, pleurae smooth.
Remarks - Phacops gran ulatus (Mun st er) was first recorded in
Poland by G. Gurich (1896) f rom the green limestones of Psiarnia containing a fa una with Posidonia ve nusta, under the name of Phacops posido niae Gtirich. That author placed it in the zone of Clymenia humboldti (Pro lobites beds).
FAMENNIAN PHACOPIDAE F R OM HOLY CROSS MTS.
127
R. & E. Richter (1926) have described Phacops granulatus from Germany, assigning them to horizons wi th Manticoceras, Prolobites and Gonioclymenia (Clym enia laevigata and Wocklumeria).
Polish specimens do no t di splay any cardinal differences as compared with those described by R. & E. Richter. The only difference is that
in the r atio of the eye length to it s d istance from posterior f ur row : in
specimens des cri bed by R. & E . Richter t his is below 3.6, while in those
of the present writer it is up to 6.4 , averagin g 4.2. Thus, while in German
spe cim ens this di stance is approximately half, the eye length, in Polish
sp ecimens it ranges f rom one third to one s ixth of the ey e length. This
may sugges t t hat P olish for ms represent an earlier s tage of the eye
migration towards th e an terior margin of the shield. This migration is
obse r va ble in the Ph acops granula tus group consisting of three species:
Phacops circumspectans P aeckelmann, Ph. granulatus (MUnster) and Ph.
wede kin di R. & E. Richter (R. & E. Richter, 1926). In the Polish specimens
of Phacops granulatus the eye is placed as in Ph. circumspectans - near
the posterior furrow with the remaining features typical of Ph .
granu latus .
Z. A . Maksimova (1953) when describing Phacops granu latus from
the Ural h as observed that these specimens have larger dimensions than
t he German forms. The latter have the cephalon 8 to 8.5 mm long, those
from the Ural 14 to 17 mm, the angle of dorsal f ur rows being 66-70 ° and
67- 75° respectively . In Polish specimens these dimensions vary strongly,
th e cephalon length being fro m 5.0 to 18.5 mm w ith the dorsal furrow
angle at 65-75 °.
Ph acops wedekindi wedek indi R. & E. Richter. 1926
(pl . I , fi g. 2 a- b )
. 1926. Phacops (P h acops) w edek i n d i R. & E. Ri chter ; R. & E. Richter, Die Trilobiten...,
p . 145, p l. 8 fig. 40- 43.
Material. - 1 incomplete cephalon from Kowala limestones.
Dimen sions (in mm) :
I. G.
169.11.6
Length of cephalon
1/2 of cephalic width
Length of glabella
1/2 of glabellar width
Width of preoccipit al ri ng (L ,) (sag.)
Length of eye
Distance of ey e fr om posterior bo rder fu r r ow
8.0
8.5
7.0
5.0
1.0
2.0
2.0
HALSZKA OSM6LSKA
1 28
Remark s. - Comparing with specimens described by R. & E. Richter
(1926) th e Polish representatives of this species d isplays the following
diff erenc es:
I
Phacops we dekindi wedekindi
I
fr om R onn e
(G erm an y)
f r o m Kowa la
(P ola nd)
2.8
2.0
2.0
2.0
L en gt h of eye (rnm )
D istance of eye Irorn p oste r io r bord er fu r row (m m)
R at io of eye length to eye di st ance f rom p oste r ior
borde r f urrow
Num b er o f lenses
G r anulati on
I
I
I
lA
1~
31
21
(2. 3, 5, 6,6.3) (2, 3. ~,4: 4, 3, 1)
?'
conspicuous
rrussing
1
A n ew su bs pecies Pha cops wedekindi uralicus Maks. ha s been d escribed f rom t h e Ural by Maksimova (1955) . The Kowala specimen differs
from it in considerably larger ey es and absence of gra nulation . This latter
fe at ure may , however, be du e to unsatisfactory sta te of preservation.
Genus T rim erocephalus McCoy , 1849
T'rune roce plui lus caecus (G u r ich, 1896)
(fig. 3 ; pI. II , fig. 1 a-c, 2 a -b . 3)
1896. Phacops caecus G iir ich; G . Giirich, D a s Palaozo icum..., p . 362, pl. Hi, fi g. 4 a-c.
1926. Ph acops (T rime roce p h allls ) caecus (Gii rich); R. & E. R ich te r , D ie Tril ob ite n ....
p . 180, pI. 10, fig . 80-8 6 (with previous synonymy).
Material. - 29 cephalons wi t h exoskeleto ns, 4 m oulds of cephalons,
4 pygi d ia wi th exoskeleto ns, 6 thorac es w ith exosk eletons - from the
Cheilocera s beds of K adzielnia (in Kielce); 1 cephalon with exosk el eto n,
2 complete specimen s enro lled, with exos kele to ns, 6 moulds of complete
s pec imens - f ro m the Cl ym en ia be ds of Kadzielnia . All the Kadzielnia
specim ens h av e be en yielde d by yellowi sh calca reous -clay sh ale, with
exoskeletons pr eserved in most cases, bu t s t rongly deformed. 14 mould s
of ceph alo ns, 1 pygidium w ith exoskeleton f rom Cl ym en ia b ed s of Lagow ,
a ll fro m dark-grey limeston e. Abou t 20 complet e specimens, en rolled,
come fro m t h e Ch eil ocera s b ed s in t he bricky ard at K ielce. These specim ens a re a pa rt of J . Czarne ck i's collect io ns . During the Warsaw insurrec-
FAMENNIAN PHACOPIDAE F R OM HOLY CROSS MT S .
129
ti on they ha ve b een damaged by fire and are thus not fit to be w orked
o ut in detail,
Dimensions of 3 specimens (in mm) :
Specimens
-
-
-+
Le n gth of ce phalon
Width of cephal on
Length of gl abella
Width of gl abella
Width of preoccipita l ri ng
(L
J)
(sag.)
Width of border a t th e po ster ior cheek angle
Dorsal furrow angl e
Length of p ygidiurn
Width of pygidium
Length of rhachis
Wid th of rh achi s
I. G.
I169.n.10
I
1. G.
I 169.11.8
I. G.
169.11.9
4.4
2.1
3.0
5.5
8.8
4.0
5.9
9.0
15.6
8.0
9.0
0.2
0.3
1.0
0.3
68°
1.0
68°
2.0
6.1
1.8
1.8
2.0
68"
2.4
Description. - Outline of cephalon sub-semicircular. Length/width
ra tio 0.56. In longi tudin al section occipital r ing high, occipital furrow
cons pic uous, preoccipital ring (L )) discernible as a small ridge between
occ ipital r ing and gla be lla. Glabell a flat , ov erhanging a nte r iorl y, in
transverse section fo rming a fl attened arch. At inner genal a ng le dorsal
fu r rows parallel, di verging anteriorly a t an angle from 70 0 to 80 0 ; occipital ring wide (sag. ), ab r up tly n arrowing a t dorsal furrows ; occip it al furrow
st rongly c urving anter iorly in m iddle par t , an d forming a shallow depression at the dorsal furrow ; p r eoccipital ring (L 1 ) a t dorsal f ur rows not
much broad er than medially , fl at t ened out , oft en with t wo tubercles
sym m et r icall y placed on eithe r side of the m idline ; p reocci pital furro w
(51) distinct, also curving for ward; second (52) and thi r d (5 3) lateral
glabcllar f urro ws r educed to two sho r t depressions , 5 3 however sometimes
m issing; ch eeks st rong ly va ult ed; lateral border broad est at th e posterior
genal angle, narrowing; anterior border situated under th e glabella is
c re scen t-like , occasionally with a sh allow concavity, sub- pa rallel to the
out lin o of t he cep ha lo n ; subcr an id ia l furrow deep and moderat ely wide;
facial sut ure detectable from mid-length of lat eral bord er follows
a course typical of this sp ecies without crossing the cheek. In the mould
the cheek is modified at its anterior angle, due to the course of the facial
s ut ure which, along a certain length , touches here the cheek, truncating
it squarely or semicircularly along the border furrow. Width of
Acta Palaeon t ol ogl ca P olonl ca -
vol. IIIj 2
9
130
HALSZKA OSM6LSKA
rhachis/width of thorax pleurae ratio 0.52. In transverse section rhachis
is high, pleurae strongly curving downward. Pygidium short and stout.
with length/width ratio 0.33. In transverse section rhachis lower than
on thorax, dorsal furrows more shallow, pleurae slightly vaulted. Rhachis ,
bluntly terminated, does not reach posterior margin of pygidium.
2c /
. ..
..
~(~j
Fig. 3. - T rimer oeep ha lu s eaeeu s (Gu r ich) : Ia cep h a lo n , dorsal view; Ib same
sp ecime n, lateral vi ew; I c same specime n, a nter ior view (I. G. 169. II. 8);
2a inter na l m ou ld of enrolled specim en, d or sa l v iew ; 2b sa m e sp eci m e n , lat eral view; 2c same specimen, anterior v ie w (I. G. 169. II. 24); 3a pygidium,
latera l v ie w ; 3b same sp e cime n, p osterior v ie w (I. G. 169. II. 25).
Ornamentation cons ist s of two k inds of tubercles: some are larger,
flat tened out and punctured, others sm all er, dispersed among the larger.
Rem arks. - Trimerocephalus caecus (Gilrich) was fo r the first time
de scr ib ed in Poland by Gil rich (1896) , f rom K adzielnia , on specimens
fr om lower Clymenia beds (zone of Cl . humboldti). In 1901 F. Dreverm ann described this spe cie s from L angenaubach (Rhine Province) and
in 19H1 - R. & E. Richter from Gattendorf. The specim ens of Drevermann a nd R. & E. Richter were found in lower Cheiloceras beds, but
accord ing to the last mentioned authors , this form may also be encountered wi thin Prolob ites b eds.
Our specimens perfectly agree with Gilrich's description. The only
ascertained difference consists in th e absence of tubercle on the occipital
ring.
131
FAMENNIAN PHACOPIDAE FROM HOLY CROSS MTS.
German specimens from Langenaubach (R. & E. Richter, 1926, pl. 10,
fig . 83) differ from those found in the Holy Cross Mts. not only in the
presence of tubercle on the occipital segment , but also in a broader.
semicircular outline of cephalon.
Trimero cephalus mastophtalmus (Reinh. Richter, 1856)
emend. R. & E . Richter, 1926
(fig . 4; pl . I , fig . 3 a - b)
1926. Phacop s ' (Trim er ocephalus) ma st ophtalm u s (Re in h . Ri chter) ; R. & E. Rich ter,
Die Tril obiten..., p. 172, pl. 9, fig. 68-74; pl. 10, fi g. 75 (with previous sy n on ym y).
1955. Phacop s (T rime roc ep ha lus ) mast ophtalmu s (Reinh . Ri chter); Z. A . Maksimova,
Trilobity ..., p. 155, pl . 14 fi g. 1-9.
Material. - 4 fragments of cephalons with exoskeletons from Clymenia beds of Kadzielnia , 2 cephalons from Clymenia beds of Lagow, one
of them as a mould.
Dimensions (in rom) :
1. G . .
169.II.7
Length of cephal on
Width of cep halon
Length of gl abella
Width of gla bella
Wid th of pre occipital ring (Lj) (sa g.)
Width of b order at p osterior ge n al a ngle
Angle of dors al f urr ow s
10.5
19.0
8.4
10.4
0.9
2.0
68 °
Description. - Outline of cephalon sem icir cular . Gl ab ell a sligh tl y
overhanging th e anterior border; its transv erse section is slightly arched,
cheeks strongly vaulted , border flat. Ma ximum width of glabella before
its mid-length; at this point gl abella nearly twic e as broad as at the
base; dorsal furrow s straight, diverging at a n angle of 68°; occipital ring
broad, strongly n arrowing at dorsal f ur rows ; occipital furrow curving
forward , de eper at dorsal furrows ; preoccipital ring (L 1 ) broad mesially,
narrowing at sides and then becoming inflated at the dorsal furrows;
preoccipital furrow (5 1) curving forward ; border broadest at posterior
genal angle, running as a uniform band as far as anterior gl abellar angle;
subcranidial furrow deep and narrow; doublure semicircular; hypostome
not preserved. Facial suture only partly discernible: it cuts the lateral
border in half of its length, slightly obliquely forward, being deeper
incised when running on the anterior part of the border; cuts the border
furrow and runs along the cheek in a concave arch, reaching the dorsal
furrow near the anterior glabellar angle.
HALSZKA OSM6LSKA
132
1
"'m
I
,
Fi g. 4. - Trimero cep }wLu s ma st opht aLmu s (Rein h. Richter) : a cephalon, lateral v iew ; b same specim en, a n terior v iew (1. G . 169.
II. 7).
Ornamentation of cephalon consists of minute gr an ulation un iformly
and fa irly densely covering the glabella and cheeks . The gran ules a r e
0.3 nun in diameter with cep halon 10 mm in length . There a re abo ut
36 granules over 1 square mm.
Rem ark s. - The here described specimens differ from those of Germany in the following characters:
Central (S,) and anterior
ral glabellar fu rrow
U:; ~) late-
from Po land
from Germany
TrimerocephaLus mastophtaLmus
·1
I d 1lsce I'm·b le
· 1 a b sen
distinct
faintly in d icated
as an uni form rid ge
narrowing at sides and
Inflated at the dorsal
furrows
Subcranidial furrow
broad
narrow
Co urse of faci al suture
di scernible througho ut
th e lateral border
discernible from midwidth (tr.) of lateral
border, deeply incised
at border furrow
Preoccipital fu rrow (SI )
I
Preoccipital r in g (L ,)
·1
I
Specimens described by Maksimova from Kazakhstan (USSR), ac cord ing to t ha t au thor , ag ree w ith t hose from German y.
Salt er (1864- 1883, p. 16, pl . 1, fig. 5-7) has figured Trimero cephalus
laevis (Munst er) wi th char acteristic glabella, st rongly wi dening anteriorly
FAMENNIAN PHACOPIDAE FROM HOLY CROSS M TS .
133
and missing second (52) and t hird (5 3) lateral glabellar furrows; preoccipital ring (L,) having large tubercles, t he cheeks bein g t riangular, the
bord er n ar row .
R. & E . Richter (1926, p . 172, pi. 9, fig. 68- 74) h av e accepted the name
of Trimerocephalus laevis as a synonym of Trim. mastophtalmus (Munster) .
Nevertheless some differences exist b etween these two forms , namely
Trim . mastophtalmus
has v ery distinct second (5 2) and third (5 3) lateral
,
glabellar f urrows , distinct facial sutur e crossing the cheek (Salter did
not m en tion the su ture in his d escription and did not figure it), while
the gran ulation on t he preoccipital ring (L ,) is very faint . Salter was
not ab le to detect .the extremely fin e granulation, characteristic of Trim.
mastophtalm us , probab ly owing to lack of ad equ at e magnifications. These
differences, however, seem to be small an d to show that w e are dealing
here with intraspecific variation only .
Specim en s of T rimeroce phalus mastophtalm u s from Poland, d escribed
in this pa pe r, come clos est to t hos e f rom Britain.
T rimeroce phalus polonicus n. sp.
(fig . 5; pl. III, fig . 1, 2 a-c)
H ol ot y pu s: I. G. 169 .11.1'2, cep h alon, fi g. 5 a n d p l. III, fig . 2' a- c.
Stratum ty picum: Famennian C l ymen ia beds.
L ocu s typ icu s: Kadzielnia within the H oly Cross M ts . area, P oland.
Deriva t i o n omin i s: p ol on i cu s - describ ed fr om P oland.
Material. - 6 cephalons wi th exos ke letons preserved, two damaged ,
all f rom the Cl ymenia beds of Kadzielnia.
Dim ension s of 2 specimens (in mm) :
Specimens
--
-
-+
Le ngth of cephalon
Width of cephalon
Le n gt h of glabella
Width of glabella
Width of preocc ip it al r in g (L1 ) (sag.)
Wid th of b or de r at posterior genal
an gle
An gle of do r sal fu r r ows
I. G.
169 .11.12
4.9
7.0
3.6
4.5
0.9
I. G .
169.n.13
4.0
6.0
3.0
4.0
1.0
0.5
59°
Diag nosis. - Outlin e of cephalon. s ub-sem icir cula r , strongly pointed
anteriorly. Do rsal furro ws slightly concave outside, their angle varies
134
HALSZKA OSM6LSKA
markedly, ranging from 51° to 80 °. Elongated dep r ess ion on the cresce ntshaped ant erior border. Facial suture crosses the cheek near its anterior
a ngle. Orn am entation of ce phalon consists of t ubercles, rather large in
r elation to cephalic dimensions, sca r cily dispersed. Two tubercles on sides
of preoccipital ring. Thorax a nd pygidium unknown.
Descri ption. - Outline of cephalon sub-sem icir cular, strongly pointed
a nte r iorl y . Length/width ratio 0.64. In longitudinal section occipital ring
high; occipital f ur row mod erately d eep ; preoccipit al ring (L 1 ) slig h tly
convex; preoccipital furro w (Sl ) m or e shallow than the occipital; glabella
somewhat ove r h an gin g th e an te r ior bord er ; border under glabella broad,
centrally d epressed. In transv erse section gl abella an d cheeks strongly
vaulted, border narrow a nd convex. Outline of glabell a st ron gly pointed
a nt er iorl y , hyperbolic; maximum glabella r width a t mid-length of glabella
b
3
mm
F ig . 5. - T r imeTocep h a lu s p o l on i cu s n. sp .: a h ol oty pe, ce p h a lon , do rsa l
view; b sa me specimen, lateral view ; c sa me specimen, an te r ior view
(I. G. 169. II. 12).
or ve r y n ear to it , a t t hat poin t gl ab ellar width more than tw ice that a t
the b ase ; le n gt h/wid th r atio 0.8; dorsal f ur rows convex ins ide, a ngle at
w hich they m eet varies strongly, ranging fro m 51° to 80°; occipit al ring
broad , na r rowing at dor sal fu rrows; occipital f ur row moderately d eep ,
d eep er near dorsal furrows, its me dian p ar t curving for war d ; preoccipital
segment (L 1 ) gently convex , wit h uniform width throu ghout a nd with ends
direc t ed post eriorly; preoccipital furrow (Sl) medially sh allow but distinct ,
mesially strongly curving forward , near dorsal furrows incised more
deeply and thus prod ucing pits directed obliquely anteriorly; second (S2)
a nd third (S3) la teral furrows m issing ; cheeks d ir ected mo r e or less vertically dow n w ard and thus imparting a character istically conical ap pe aran ce
tv the w hole cephalon. P ost er ior bo r de r narrow a n d con vex , broadening
a nd fl attening out wh en passing into lateral border wh ich here ' a tt ains
its maximum width. Viewed fro m above t he border is discernible nearly
a s fa r a s the lat eral angl e of glabella ; it t he n pa sses under the glabella
FAME N NI AN P HACOPIDAE FROM H O LY CROSS MTS.
135
where it is broad and cr escen t-sha ped ; here an elongate, wide depression
occurs separating the anterior border into two convex lists; subcranidial
furrow deep and wi de; border furrow deep, highly arched under the
gla be lla . Eyes mi ssing. Facial sut ure ve ry faint, discermble on the cheek
on ly wher e it is shape d like a sma ll ar ch and enters the dorsal fu r row in
f ro nt of the gena l a ngle.
Ornam en tation of cephalon ve ry characte r istic: glabella and ch eeks
with t ube rcles from 0.07 to 0.1 mm in diamet er (the cephalic length being
4.4 mm, its wi dt h 6.5 mm) . The t ube r cula t ion is distinctly spa rs e, th ere
being about 16 granules on ea ch mm . On the preoccipital ring (L I ) two
t ubercles ar e pl aced symmetrically on either side of the centre , not larger
than the tubercl es on the glabella a nd cheeks. Occipital ring and border
s moot h. In fro ntal part of glab ella and on border under the gl abella ,
tub erculation more dense.
R em ar k s. - This species do ubtlessly belongs to genus Trimeroc epha lus , as is s uggested by such characters as: fa lcate shape of border below
g lab ell a , facial sutu re crossing th e cheek , preoccipital furrow unb roken
in the m id dle part and curving forward . On the whole this species most
a pproach es Trimerocephalus caecu s, its glabella , however, is more sharply
po in ted and th e course of its facial s ut ure differs in that it not only touches
the cheek basall y but even crosses it. A diagnostic feature here is the
differ ence of ornamentation . In Trimerocephalus polonicus n. sp. it consists of fa irly large t uber cles spa rsely dispersed on the gla bella and
cheeks.
In Trim eroceph alus m astophtalm us (Re inh. Richter) , described by
R. & E. Richt er (1926, p . 172, pl. 9, fig. 68-74) , sim ila rl y as in Trim. polonicus, the cheek is t r an sected by th e fac ia l sut ure , while the dorsal f ur rows
curve ou ts ide. In th e former, howev er , the outline of cephalon is sem icirc ular , gl ab ell a sli ghtly rounded anteriorly wit h di scernibl e a n teri or
a nd cent ra l later al furro ws a bsen t from the her e d escrib ed species. Ornament ation of ceph alon in Trim. mast ophtalmus con sist s of fine, d ens e
granulation, th e length of cephalon ra ngin g fr om 5.2 to 16.5 mm, while t he
largest cephalons in Trim . polonicus n ever exceed 4.4 mm in length and
th eir ornamenta tion con sist s of markedly sparsely dispersed granu les.
Genus Dianops R. & E . Rich ter , 1923
Dianops typhlops (G iirich , 1896)
(p l. IV , fig. 1, 4)
1896. Trimerocepha!us t y ph !o ps Gtirich ; G. Gtirich, Da s Palaozoicum..., p . 359, pl . 115,
fi g. 7 a b.
1926. Phacops (T r i mer ocep ha!u s) typh!ops (G tirich); R. & E. R ich ter , Die Trilobi ten...,
p . 192, pl. ,10, fig. 90-94 (with previous syn on ymy ).
136
HALSZKA OSM6 LSKA
Mate r ial. - 4 cep halons, 2 fragments of pygidi a, wi th exosk eleto n
pa rtly pr eserved, from dark-grey Clymenia limestones of Her by.
Dim en sion s of 2 specimen s (in mm):
Specimens Leng th of cephal on
Wid th of cephal on
Length of glabella
Wi dt h of glabella
Length of pyg idium
Wid th of pygidiu m
Len gth of rhac h is
Width of r h achis
-+
1. G .
I 169.II .14
I
I. G.
169.II .15
8.0
16.0
7.0
8.6
2.3
5.0
2.1
1.2
Rem ark s. - Dianop s typh lops (Giiri ch) h as been descr ibed fro m P oland i. a. also by G u r ich (1896) a nd R. & E. Richter (1926). Upon comparison with t he lect oty p e (R. & E . Richter, 1926) t he h ere d escribed
specimens d isplay some sligh t diff erenc e. In t he longitudinal se ction ,
namely , t he ce phalo n is less v aulted , wh ile the lateral border is relatively na r row er.
Dianops? trifo lius n. sp ,
(fig. 6 ; p l. IV, fi g. 2, 3)
Holotypus: 1. G. 169.II.16, ce p h a lon, fig . 6 and pl. IV, f ig . 2.
Stratum typicum: F amennian Ch eiloceras bed s.
Locu s typicus: Kadz ielnia w it hi n the H oly Cross M ts . are a , P oland.
D er i v at i o no m i n i s: -trifo!ius - with sh ape r es embling a trilobate leaf of clo ve r
(T r ifolium) .
Mat erial. - 5 cephalons, with exoskeletons pr es erved, fr om the Cheil ocera s be ds ; 2 d amaged cephalons, w it h e xosk eletons, from the Cl ymenia
beds of Kadzi elnia. Nearly all the specimen s sli gh tly d amaged.
Dimen sions of 3 s pecimens (in mm) - s ee p . 137.
Diagnosis. - Outlin e of cephalon trilobate. Gl ab ella flattened anteriorly, t w ice as broad as at t he base, lateral angles gently truncated. At
posterior ge nal angle the border twice as broad as at the anterior angle,.
bent inside t he dorsal furrows. Thorax and pygidium unknown .
Des cript ion. - Outline of cephalon that of a semicircle strongly bent
inside .the dorsal furrows. Length/width ratio 0.5. In longitudinal section
occipital ring lower or even as high. as glabella; .preoccipital segment (L 1 ) ,
F AM E N N I AN PHA COPIDAE FR O M H O LY CROSS MTS .
Spe cimens
-- -~
L engt h of cephalon
1/2 of cephal on wi d th
Length of gl a b ella
W~ dth of glabelJ~ . .
Width of
W id th of
an gle
Wid t h of
a ngle
An gl e of
p r eocc ip ital r ing ( I. , ) Isag.]
borde r at po st e rior ge n a l
I
I
I
137
1. G .
169.II .16
I. G .
169.lI.l 8
1. G .
169.lI.1 7
4.5
5.0
3?
5. 1
6.0
5. 1
5.0
6.2
0.4
6.0
6.5
5.0
0.5
0.9
1.5
borde r at a n te r io r ge n a l
d or s al fu rrow s
0.8
70"
0.5
70°
0.8
68"
only occasion ally prom in ently va ulted ; preoccipita l fur ro w (S,) faintly
indicat ed ; gl ab ella is r aised , ge ntly a rched, not ov erhan ging the bo r-der ,
with a small front al edge; borde r w eak ly va ult ed . In transverse section
glabella an d ch eeks gentl y arcua t e, border wide , mod er ately convex .
Glabella broadly ro un ded ant er ior ly, att aining its m ax imu m width in front
of m id-length ; length/widt h ratio of gla bell a 0.7; its shape r esem bling
a trapeze wi th base h alf th e length of th e upper edge; lateral ang les of
c
·.b+2";·'~·:+:R,
3
mm
Fig. 6. - D ian op s? trij ol ius n. s p.: a h olot y p e, cephalon, d orsal v iew; b sam e
specimen , la ter a l v iew ; c same specime n , a n te r ior v ie w (1. G . 169. II. 16).
gl ab ell a som ewhat tr uncated; dors al fur rows strai ght , d iverging at an
angle of 70°; occipital ring moderat ely wide, occipita l furrow st r aig ht;
preoccipital ring (L ,) narrow, pr eoccipi tal furrow (S,) stra ight, us uall y
interrupted mesially; pits in preoccipital and occipital fur row ve ry shallow ,
elongate; two anterior lateral glabellar (S2' S3) f ur rows no t discernible;
cheeks in the sh ap e of an almost equilateral triangle ; posterio r border
narrowing at mid-length (tr.), th en wid ening and att ainin g maximum
width where it meets the border; the border wide posteriorly, narrows
anteriorly so that it is there nearly 2 times narrower than at the posterior
genal angle; at contact with dorsal furrows the border strongly bent
inside, viewed dorsally it is seen along 1/3 of the length of the anterior
138
HALSZ K A OSM6LSKA
margin of glabella and farther being concealed beneath it ; under glabella ,
limbus broadly fiat and running as a uniform band; subcranidial f urrow
deep and broad, with h ind edge somewhat lower than the frontal; border
furrow deep and fairly wide, bent inside at the dorsal furrows; under
gl abella subcranidial and border furrows running parallel. Facial sut ure
no t discernible. Ey es m issing.
Ornamentation of cephalon: Glabella , ch eeks and border near the
dorsal fur row covered by minute granul es 0.03 mm in diameter, with
cephalon length 6.0 mm. Und er X 30 magnification occipital ring shows
delicate granulation . In badly preser ved specimens , particularly on glabella, granulation not visible.
Remarks. - This species displays following features suggesting its
as signment to genus Dianops: before glabella limbus broadly flat, under
glabella subcranidial and border furrows parallel; second (52) and third
(53) lateral furrows missing; preoccipital furrow (51) straight , broken in
the middle part.
Other features of this species, however, do not agree with the diagno sis of genus Dion ops, as given by R. & E. Richter (1926) . They are,
namely, different shape of glabella which ex pands uniformly forward ,
a nd anterior angles of glabella only quite insignificantly truncated. Moreov er, in our species, th e border is throughout its length distinctly delimited f rom the cheek by a deep border furrow.
Bending of the bord er at the dorsal f urrow, characteristic of Dianops?
tTifolius n. sp ., may have been caused by the exosk eleton being broken
off h er e along the facial suture, which has a sim ilar course in genus
Dianops. This, howev er, is rath er doubtful , sin ce in blind t r ilob it es t he
sut ure w as not , in all prob ab ility, f unc tional.
In comparison wi t h ot her species of gen us Dianop s, Diano ps? t rifolius
shows the following diff er en ces: outline of it s cephalon is sem icir cular,
bent n ear the dorsal furrow (this is a cha racter not en counter ed in any
one of the d esc r ib ed species of ge n us Di an op s) ; lat eral angles of gl ab ella
only sligh tly truncat ed (in Di an ops typhlops, D. an ophtalmus, D. griffit hid es glabellar ang l es are conspicuously truncated) ; the en t ire glabella
and cheeks, as well as border n ear the dorsal furrows are finely gran ulated . Granulation also occurs in D. t y ph lops, D. anophtalmus and D. griffit hides, but in some pa r ts of the cephalon only.
Phacops sp.
(fig. 7 ; pl. III , fig. 4)
Ma t erial. Kad zielni a .
1 py gid ium with exos ke le ton fo und in Cl ymenia beds of
F AM E N N I AN
P ~AC O PI DA E
139
F R OM H OLY CROSS MTS .
Dim ension s (in mm) :
I. G.
169.I1 .19
Length of pygidium
Wid th of pygidium
Length of rhachis
Width of rhachis
5.2
10.2
5.2
3.1
Description . - Width of pygidium more than twice t he length :
o utlin e of posterior margin gently arcuate, sli ghtl y concave where rhachis
r eaches the margin . In longitudinal section, rhachis considerably el evated
-over ple urae, this elevation eq ualling 2/3 of pleurae h eig ht; three distinct
segme nts and con spicuo us rha chial r ings, thre e further seg ment s not r ead-
F ig . 7. -
Pha cops sp .:
a p ygidium, dorsa l v iew; b same spe ci m e n , la tera l view ;
c same sp ecimen, po steri or view (I. G. 169. II . 19).
ily discernible; rhachis ben t rather sh a rply where it m eets posterior margi n of shie ld, r eaching to end of shield. In transverse section rhachis
s t ron gly va ulted . Dorsal fu rrows converging V-shaped at posterior m ar gin
of shield ; pleurae horizontal from dorsal furrows to approximately 1/3 of
t heir width (tr.) , farther strong ly curved downward; posterior margin
Tc.ised wh er e m eeti ng t h e rhachis; 6 furrows between rhachial rings, the
frontal ones semicir cular an d conspicuous, deeper along dorsal furrows ,
beginn ing with the 4th barely discernible and m esiall y angularly bent;
4 r ibs present on pleurae, gently arcuate distally. P le ural grooves mod erately deep, intersegmental furrows mor e s ha llow b ut likewi se distinct ;
no t one of them r eaches to m argin of shield; a nterio r and posterior po rtions of ri b dist inctly d elimit ed , conv ex ; segm en ts of r ha chis wit h very
fa int ornamentation consisting of low tubercles 0.07 rom in diameter.
Remarks.
The above described pygidium differs considerably
from pygidia of all species of Phacops so far described. It may possibly
be referred. to one of the two n ew sp ecies described in the present paper
found in the same beds and presenting a similar fine granulation.
HALSZKA OSM6LSKA
140
Subfamily Phacopidellinae Del o, 1935
Genus Du ctina (R. & E. Ri cht er, 1931)
Ductina du ctifrons R. & E . Ri cht er , 1923
(pl. III, fig. 3)
1926. P hacopidella duct ifron s R. & E. Ri ch t er ; R. & E . Ri ch t er, Di e T rilobiten...,.
p . 207, pl.1'l, fi g.
1 6 ~2i2
(w ith pr ev iou s syno n ym y ).
1·931. P ha eopideUa (D uetina) d u et ifrons R. & E. R ich t er ; R. & E . R ichter , U nter 1a gen
zum Fossilium Ca tal ogu s.
19515. P h aeopi d eUa (D uetina) du et i f r ons R. & E. R ichter ; Z. A. Maks imova, T r il ob ity ..,
p. 1186., pl. 1,6, fig. 1-6 .
Mat e'riaL - 4 cep halons from Clym en ia bed s of P siarnia. These sp ecimens, wi th exos ke letons , ha ve been pr es erved in shal es, in a d eformed
and fl attened con dition.
Dim en sions of 3 specim ens (in mm):
S pe ci m en s - - -,
Length of ce pha lo n
Wi dth .of cepha lon
L en gt h of glabell a
Width of g labella
An gl e of d orsa l f ur r ows
1. G .
I 169.11.20
I 1. G. I 1. G.
1169.11.21 169.11.22
4.0
7.1
3.1
I
5.0
4.1
4.1
60 0
600
Descri ption. - Outline of cephalon semicir cula r, le ngth/width r atio
0.5; dors al f urrows divergent at an angle of 58° to 60° , fro ntally curvin g
to t h e outs ide and passing un int errupt ed in to borde r furrow ; gl ab ell a
widening o ut a nteriorly, wi th late ra l angles passing outside into the
border ; len gth/wid th ratio 0.8; occi pital ri n g m oderately narrow, pr eoccipit al ri ng (L 1) b ro adly fl at , occipit al a nd preoccipital f urrows (S1)
delimiting it straight ; preoccipit al fu rrow (S1 ) m edi ally v er y fain tly
indicated; eyes a bsent .
'R em ark s. - P hacopid ella ductifr ons (1926), described by R. & E. Ri chte r , is wit ho ut s ubcran idia l fu r row, w hi ch is a gen eric character, while
th e fa cia l sut ure r uns along th e out er m argin of border. Exoskeleton with
delicate granulat ion. The he r e m en tion ed features a re not discernibl e in
Polish forms owing to ext r emely bad sta te of preservation. Ductina
duc tifrons has not thus far been d escribed from Poland. In Europe it has
been recorded from the Rhine Province and from Eastern Thuringia, as
well as from Great Britain. After R. & E. Richter (1926) it is encountered
within Mant icoceras and Cheiloceras beds.
Paleozooloqical Laborat ory
of the P orish Academy of Sciences
Warszawa, January 1958
FAMEN NIAN P HACOPIDAE FROM H O LY CROSS MTS .
141
REFER ENCES
·C ZA RN OCK I, J . 1928. Przeglad stratygrafii famenu i k arbonu dolnego w zachod niej i sr odk ow e j czesci Gor Swietokrzyskich. - Pas. Nauk , P . In st. Ge o!.,
21, 55-61 , Warsz awa.
1948. Przewodnik XX zjazd u Polskiego T ow a r zy s tw a Geol ogi czn e go w Go rach Swietokr zysktch w '1l»7 r. - Roc zn. P . T. Geo!. (Ann . S oc. Geol, de Pologn e), (1947) 17, 237-299, K r a k ow .
Ii!};}7. Geologia re gi onu Iysogorskiego (Geology of the Lysogory Region). Prace Geo!., 2, 3, 34-35, Warszawa .
DE LO, M. D. 1940. Phacop id Trilobites of North America . - Ge o!. Soc. Amer.,
S pec. Pa per s, 29, p . I;}, New York.
DR EVERMANN, F . 1901. Di e Fauna del' oberdevonischen Tuffbreccie von Landenau bach be i H aiger. - Jb. preu ss. geo!. Landesan st ., (1900) , p . 114, Berlin.
G DR I CH , G. 1896. D as P ala ozoi cum im po lnisch e Mittelgebirge. - Verh . Russ.
K ais. Min . Ges., 2, 32, 35'9-356, St. Petersburg.
HUPE, P . 195'3-19515 . Classification des Trilobites. - Ann. Paleont. , 39, 41, 249-263,
P a r is.
JAANUSSON, V. W5i6. On the Trilobite ge n u s Celmus Angelin , 1854. - B u l l , Geol.
In st. Uni v. Upp sala, 36, 36-37 , Uppsala.
MAKSIMOVA, Z. A. 1955. Trilobity srednego i verchnego devona U r a la i severnych Mugodzar. 136-'142, 155 -160, 186-188, Moskva.
RI CH T ER, R. & E. [ 919. Uber Phacops laevis und andere P hacopiden des vogtlandischen Oberdevons. - Senckenbergiana, I , 4, 131-'1'40, Frankfurt a . M.
192.3. Uber Phacopidella Ree d . - I bidem, 5, 3/4, 134-142.
1926. Die Trilobiten de s Oberdev ons. - A bh . preuss. geo!. Landesanst. , N . F .,
99, '112£- 211, 236-245, Berlin.
1931. Unterla gen zum Fo ssilium Catal ogus, Trilobitae: 5. - Senckenbergiana ,
13, 2', F r a nk fu r t a . M.
ROE MER , C. F . 1186,6 . Geognos ti s che Be ob a ch tu n gen im P oln is ch e n Mittelgebirge. Ztschr; deutsch . geo!. Ges., 18, Berlin.
SALTER , J . W. 11864. A Mo no graph of British T r il ob ites. - Mo nogr. Pal eontogr.
Soc., 1'6-'17, L ondon.
SAMSONOWICZ, J . 1917. U tw or y dew on skie w s ch od n ie] czes ci Gor Swietokrzyskich. - Prace T ow . Na u k: Warsz., III Wydz., 20, Warsza w a ,
S OBOLEV, D. 19111. 0 fa m e nsko m jaruse Kelec ko -sandornerskogo krja za . - Ezei) .
geol. miner. Rossii, 13, 1/2 , 34-4 1, Novo-Ale ksand r ia.
H ALS ZK A OSM6LS KA
PHACOPIDAE F AMENSKIE Z GOR SWI.E;TOKRZY SKICH
Streszczenie
P h acopidae f amenskie,
opisa ne w
niniejsze]
pracy,
zostaly
zeb r a n e
prze z
Dr Z. K ie la n w nastepujacych miejscowosciach: K adzielnia , Psiarnia , H erby (dzielnice Ki e lc), oraz Lagow , Ga lezice , Kowa la i Ja blonn a ko lo Borkowa (fig. 1 ) i prze-
HALSZKA OSM6LSKA
142
kazane arutorce d o opracow a nia . Wszystkie te p un k ty po lozo n e sa n a p oludn ie od
nasuniecia Sw ietokrzy sk iego i zgrupowane w dw och odr eb nych fauni stycznie i litologiczriie regionach: I. synklinorium cen tr al ne (kielecko-la gowskie), II. synklina
galezicko-daleszycka.
Famen w synklinorium cen tra lnym w yk szta lcon y jest w po st a ci ponad 100-metrowej serii osadow lupkowo-wapiennych
0
szaro-zoltawym zab a r w ie n iu.
Fauna
trylobi t owa jest d os e obfita; sa to w ylac zn ie trylobity slep e z ro d zin y Phacopidae :
Trimerocepha lu s caecus (Giirich), Trim. mas t ophta lmus (Reinh. Ri chter), Trim .
polonicu s n. sp .,
D ian ops t y ph l ops (Gu r ich ),
Dianop s? trif ol ius n. sp. ,
D u ct i n a
ductifr on s R. & E. Ri chter. Charaktery st yczny tu jest zupelny brak trylobitow z ro-
dziny P r oe ti d ae. Osady t ego typu wyksztalcon e sa w nastepuj a cych punktach:
Famen sp oczy wa tu n a w apieniach franu ; sa t o lupki rna rgliste-
Kadzi elnia -
przewar stwiane w ap ien iam i. T worza one gruby k om p lek s z fauna trylobi tow i m alZOW oraz zle za cho wanych klimenii. Wedlug J . Czarn ockiego (J.9~8 ) sa tu reprezento w a ne w a rs twy ch eiloc erasow e i klimeniow e. Sposrod Phacopidae wystepu ja: Tr imer ocep halus caecus (Giirich), T rim . ma st ophtalmus (Reinh. Ri chter), Trim. polon i cu s n . sp ., D ia n op s? trifolius n. sp .
Ps ia rnia -
F amen je st tu wyksztalcony rown ie z w postaci lupkow marglistych
przewarstwion y ch wapieniami. Phacopidae r eprezentowane sa przez : Phacops granu l at us (Mu nster ), D i an op s typhlops (Giirich), Ductina ductif r ons (R. & E. Ri chter ).
W drodze prowadzqcej do Czarnowa odslaniaja sie czar ne wapienie;
Herby -
w ktorych spotyka sie Dianops t y p h lo ps (Giirich).
Lagow
Od stoniecia famenu wystepuja na polnocnym krancu m iastaczka,
w zboczu doliny rzeczki Lagowicy. Sa to silnie bitumiczne osady lupkowo-wapienne.
W ciemnoszarych w ap ieniac h klimeniowych znajduje sie niekiedy wielkie n a gr c madzenie p a n cerzy Trimerocephalu s ca ecus (Giirich) i nieliczne Trim. mastophtalmus:
(R einh. Ri chter). Z innych Phacopidae cytow ano tu Dianops typhl op s (Giirich). Wy ksztalcen ie fame nu w Lagow ie odbiega ni eco od typowego lupkowo -wapiennego,.
gdyz pr zew aza ja tu wap ie n ie, Iu pkow zas jest mniej . Zesp ol faunistyczny jest tu
tak ze odm ie n ny :
Proetidae.
oprocz bar dzo li czn y ch Phac op id a e, sa tu trylob it y z r odziny
F amen w synklinie ga le zicko - da leszy ck ie] w yk sztalcony jest w f acji wapiennej .
Sq to wapien ie szar e, czest o
0
zabarwi eniu czer w onym,
0
miazszosci nie przekra-
czajace j 30 m . Fauna ich jest bardzo bo ga ta i d obrze zachowana. Sposrod Phacopidae w ys tepuj a tu tylko dwa gatunki; sa to formy majace oczy : Phacops granulatus (Muns ter) i Ph . wedekind i wede k indi R. & E. Richter; natomiast Proetldae ,
ktorych niemal zupelnie brak w synklinorium centralnym, sa tu bardzo Iiczne,
Osa dy fameilskie typu wapiennego wystepu ja w nastepujacych punktach:
Ga!~zice
-
Br a k tu fran u, famen zas spoczywa b ez posrednio i p raw ie zgodnie
na wapieniach amfiporowych zywetu. J est on wyks ztalcony w p osta ci twardyc h.
wapieni
0
zabarwieniu cie mnosza rym , rozowym, czekola d owym i zielo nkawym. We -
143
F AM E NNIAN P HACOPIDAE FROM HOLY CROSS MTS .
drug Czarnockiego (11928) brak tu poziomu cheilocerasowego, a gorny famen obejmuje ws zystkie poziomy klimeniowe, Iacznie z woklumeriowym oraz gattendorfiow ym , nalezacym juz do dolnego karbonu. Fauna jest b ardzo bogata i doskonale
zachowana ; zawiera on a goniatyty, klimenie, or tocer asy, brachiopody , m alze, trylobity, s zcza tk i ryb. Sp osrod Phacopidae wystepuje Phacops granulatus (MUnster).
Wskutek w yk li n ow a n ia sie pewnych w a r s tw miazszosc w poszczegolnyeh odkrywk ach w sarnych Galezicach waha sie od 3 do 7 m.
Kowala -
Famen wyksz talc ony tu jest takze w po staci w apieni, czest o czer-
wo n o zabarwionych , gruzlowaty ch , le za cy ch na w apieniach franu. Wedlug Czarn ockiego C1957) w y s tep u ja tu ws zys t k ie poziomy klimeni ow e. Sposrod Phacopidae w y-
step uja:
Dian ops typhlops (Gtirich) i Ph. wedek in di we de k i ndi R. &
E. Ri chter.
Wyzej Ieza lupki dolnego karbonu.
Jablonna k . B orkowa -
Famen zostal tu odkryty przez Czarnockiego p o II
wojn ie swiatowei. W ro ku 1949 pro w adzit on t u pra ce geologiczne, kt6rych wyn ik i n ie zost al y jednak opublikow ane. Famen za lega tu n a w a pie nia ch franu; jest
to , podobnie ja k i w Kowali, kompletna se r ia, za czynajaca sie od warstw cheilocerasowych i prawdopodobnie ze wszystkimi p oziomami klime niowymi. Sa to wapienie
o czerwonym, szarym woklumeriowe -
a w wa r stwach in ter p retowany ch przez Czarnockiego jako
seledynowym zabarw ieniu. F auna b ardzo bogata. Miazszosc okolo
30 m . Sposrod Phacopidae wystepuje tylk o Phacop s granulatus (Munster). Przedstawiciele te go gatunku, wystepu iacy w nizs zych poziomach klimeniowych, to osobniki
0
malych, kilkumilimetrowych cefalonach. Ten sam gatunek z seledynowych
wa p ie ni woklumeriowych osiaga rozmiary ce fa lon u dochodzace do 18 mm. Nad osadami famenu le zq lupki dolnego k arbonu.
Wsrod opis a n y ch prze z autorke Pha copidae fameriskich w wymienionych w yze ]
punktach w ys tepu ja trzy ga tu nk i, zn ane [u z przedtem z G6r Swietokrzyskich :
Phacop s granulatus (M unst er )
Trim erocephalus caecu s (Gtirich )
Dianops typhlops (Gtirich)
a takze dotychczas w Polsce nie zn ajd ow ane:
Phacop s wedeki n d i w ede ki ndi R. & E. Rich ter
T rime ro cep ha lu s mastophtalmus (Reinh. Richter )
Duct ina ductifrons (R. & E. Richter).
Do tego d ochodza dwa n owe gatunki:
Trimerocephalus p ol onicus n. sp ,
D ia n ops? trifolius n. sp.
Ponizej przytoczone sa kr6tkie diagnozy ga t un k 6w nowych.
Trimerocephalus polonicus n. sp,
(fig. '5; pi. III, fig. 1', 2 a-c)
H olotypus: I. G. '1 69. II. 12, cefalon, fig. 5 i pi. III, fig . 2 a-c.
Strat u m typicum: fa m en, war stwy klimeni owe.
HALSZKA OSM6LSKA
144
L ocu s ty picu s: K adzielnia , G6ry S w ietokrzyskte, Polska,
opisany z Polski.
D er i v a t i o n ominis: p ol on i cu s D iagno za. -
Zary s ceIa lonu w ez szy niz p 6lkole; glabella z przodu silnie za-
os trzon a; br uz dy g rz b ietowe lekko wy giete, w k les le na zewnatrz, kat ich bardzo
zmie n n y (511- 80 ° ); na
p ol k siezy cowa tym li m busie przednim podluzne wglebienie ;
sze w t war zowy prz eci n a po lic zek p rz y przedni m [ego kacie ; or nam en ta cja cefalonu
w p ostaci d ose d uzy ch w s tos u nk u d o w ymia r6 w ce fa lonu guzkow, rzadko rozmi eszczon ych: p o bo ka ch m ied zy pi erscienia d w a guzki. T oraks i pygidium ni e
z na ne.
Dianops ? trifotius n . sp .
(fig. 6 ; p l. IV, fi g. 2, 3)
H ol o typus: I. G. 169. II . 106, cefa lon , fig. 6 i pl. IV, fig . '2 .
S t ra t u m t y p icum: fa men, warstwy ch e il ocer a s ow e.
L ocus typ icu s: Kadzielnia, G6ry S wi et okrzyskie, Polska.
Derivat i o n om in is: t r ifoli u s D ia gn oza . -
ce fal on rn a ksztalt li sci a koniczyny (Trifol ium).
Zarys ce fal onu przypominajq cy tr6jplatowy lise koniczyny; gla-
b ell a z przodu spla szczona, d wa razy sze rs za niz u podstawy, jej boczne katy lekko
sci ete ; limbus p rzy tylnym kacie policzkowym przeszlo dwa razy szerszy niz przy
przedn i m, przy ujsciu bruzd grzbietowy ch wgiety do wnetrza . Toraks i pygidium
n ie znan e.
OBJAsNIENIA DO ILUSTRACJ I
Fig . 1 (p . 122)
M ap k a w ys te powa n ia famenu w p oludniowej czesci G6r Swietokrzyskich.
Fig. 2 (p . 126)
Phacop s granula tu s (Mu n s te r ): ta cefalon, profil podluzny : Ib ten sam okaz,
profil p opr ze czn y (I. G . 169. II. 1); 2a pygidiurn, profil pcdluzny ; 2b ten sam okaz,
prom p oprzeczny (1. G. 169. II. 23).
F ig . 3 (p. 130)
Tr imerocephalus cae eu s (Gil r ich ): la ce fa lon, z g6ry ; Ib tel] sam okaz, profil
po d lu zn y ; t c t en sam okaz, profil poprzeczny (1. G. 169. II. 8); 2a oka z zwiniety,
os r od k a w ewnetrz na, z gory ; 2b ten sam oka z, profil podbuzny ; 2c ten sam okaz,
profil poprzeczny (I. G. 169. II. 24); 3a pygidium, profil podluzny ; 3b ten sam okaz,
profil poprzeczny (I. G. 16'9 . II. 25).
Fig. 4 (p. 132)
(Reinh. Richter): a cefalon , profil podluzny ;
b ten sam ok a z, profil poprzeczny (I. G. 169. II. 7).
Trimerocephalus
mast ophtalmus
Fig. 5 (p . 134)
Trimeroeephalus polonieus n. sp.: a holotyp, cefalon, z gory;' b ten sam okaz ,
profil podluzny : e ten sam okaz, profil poprzeczny: (I. G. 169. II. 1(2).
FAME N NIAN PHAC O PIDAE FRO M H O L Y CROSS MTS .
145
F ig. 6 (p. 137)
D ian o p s? t rifolius n . sp. : a h olo t yp, cefalon , z gory; b ten sam okaz, profi l
p od lu zny ; c ten sam oka z, profil pop rzeczny (I. G . 100. II. 16).
Fig. 7 (p, 139)
Ph aco p s sp.: a pygidium, z gory; b ten sam ok a z, profil pod luzny ; c ten sam
okaz, p rofil po przeczny (I. G . 169. I I. 19).
P i. I
Phacop s granulatus (M u n ste r)
Fig. 1. Cefa lo n , z gory (I. G .169. I I. 1); X 5.
Phacops wed ekindi wedekindi R. & E. Richte r
F ig. 2. a Fra gment cef a lo n u, z go r y ; b ten sam ok a z, profil p odluzny (I. G. 169.
II. 6) ; X 4.
Trim er ocephalus ma st ophtalmus
(Re i n h . R ich ter )
Fig. 3. a Cefalon, z go ry; b ten sam okaz, profil poprzeczn y ; c ten sam ok a z,
profil pod lu zny (I. G . 169. II. 7) ; X 3,1.
P I. II
Tr imerocephal u s caecus (Gu r ich )
Fig. 1. a Okaz zw in iety, z gory ; b ten sam ok az, p r ofil p opr ze czny ; c ten sam
ok az , profil podhizny (I. G . 169. I I. 8) ; X 4,7.
Fig. 2: a Caly oka z, osrodka 'w e w n etr zn a , z go ry; b ten sam ok az , pygidium
(I. G . '169. II . 11) ; X 2,7.
,
Fig. 3. Wapien z lagowa, szlif m ikrosk opo wy ; w idoczne pr zekroj e p an cerzy
Trime rocephalu s caecus; X 3.
Pi. I II
Trimero cephalus poloni cu s n . sp.
F ig . 1. P ar atyp, splasz czony cef a lo n, z go ry . (I. G . 169. I I. 26) ; X 6,4 .
Fig. 2. a H o1oty p , ce fa lo n, z gory; b te n sa m ok az, profi1 p oprzeczny ; c ten sam
o ka z, p rofil p odlu zny (I. G . 169. II. 1'2); X 8,7.
Ductina ductifroll s (R. & E. Richler)
Fi g. 3. Fragment ce fa lo n u, z go ry (1. G . 169. II. 20); X 9,5.
Phacop s s p .
Fi g . 4. Pygidium , z go ry (I. G . 69. II. 19); X 4,8.
Pi. IV
D i an o p s typhlops (G urich)
Fig. 1. Cefalon, z gory (I. G. 169. II. 14) ; X 4,7.
Fig. 4 . Pygidium , z gory (I. G. 169. I I. 15) ; X 8,7.
Dian op s? trifolius n . sp ,
Fig. 2. H ol oty p , cefalon, z gory (I. G. 169. II. 116) ; X 8,4.
Fi g . 3. Pa ratyp, ce fa lo n, z gor y (I. G . 169. II. 17); X 5,6.
.Acta P al a eon t ol ogi ca Pol ont ca -
vol. III j 2
10
HALSZKA
146
OSM6k~KA
rllTIbWKII OCMYTIbCKII
cPAMEHCK11E PHACOPIDAE 113
ror-
C B E H TO K P lli 11C K 11X
(TIOJIbWA)
Pe 3lo~e
Omlc.3J!i1Hbie B ::n 'OH CT,aJTbe epalM0H CllrH 8 Phacop ida e COOp.'lIHbI )]Jp . 3 . KeJlilIH 'B cne n y ioutx M€ C'flHOCTHX : K alL\3 eJlbi HH, TICHipHIH H r ep6 bI
JI€lH3n ~e ,
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p aaoaax :
I.
~€lH11paJlbHbIH ('R eJIe~RO -JIaI'O!BCRHrr) Ol1'HiKJFlIJl-llQrHH,
II.
rlliJI eJH3M'~RO-AaJIe­
III H~l\aH 'Cll1lHiRJI:l tHJaJ] b.
$aJ~eH B ~€lHrnplliJIbiHOM CIiiHiKJI:l1IH1OpHe IlIpeACT<LBJIeH CBlf.l'OH 008JH~etBa'I10-H3IB€C'flIIIH ­
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CJI'JIeJlb HO
,M HOI' Oq !1'CJI€lHlHla,
ceMe H'C'I~Ba Ph a c op id a e
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C ord a ) :
WO
-M . llr1tJlooHwsaJH epa'Y'H13 OrJ1HO-
M'ClKJIIOqrHTeJlbl HlD ooeIlbIMH
epOpMaMH
T ri m er oc ephal u s caecus (Giirich), T r i m.
ma sto p htalmus (Rei n h. R ich t e r ) , Trim . polonicus n. sp., D i an op s typhlops (Gurich ),
D i an ops? trifolius n. sp ., D uc t i n a ductijrons R. & E . Rich t er. XClIpa'KTepHCT:Hq€OKH.YI
ep8JRTO)1 HBJIH€ TICH 3'Ae,c b
n o aaoe
01' CyTCl'BH€
11P HJlOOH'flOB H 3
P r oat ida e,
ICIffivl€ HCTIBa
OC3~RlH y;noMm~yroI'O -rnna Ip a3 B HTbI B OO€AY I<l ~X " YiHIRTaDC
Raj\3eJlbIUI a
'ro qrHOO ara
cP,a cvI€tHI:JKHH
~epreo1HCThIX
spyc
CJIaJHI~ra.x
aaneraer
snec s
aa
ITe'PeCJI@H'BaI<l ~H X('J1
<ppaJID:JKHX
C
H3IBeCTiHHKa('{.
IH3 BOO11HHlK a 1\U! .
OHH
oepasyior MOI..I.VHlbIH IKOMI1.JI€lKrC 0CaAKOB C epa Y;HoH l1PI{JIOOHTOB H ITJIaCTH:wla 11OmaOe[j llbIX Ip HiAQM C lPPaJI'M€IH11aJMIH rK ~HMe.HJH H ITJIOX OH OOOOPaJHlHOCTH.
no H.
lJ aJPifliO~KHJM (.194 8)
F.S11OM 1ll:Y1H1K1'e rnpencraane nsr XeHJIO~€lpoB ble IH I j{ JI~lM>eHIHes Lle CJI0H. 113 qrHCJHl> Ph aco pida e ornpeA€1JI€IHbI : T r imeroceph al us caecus
(G iiric h ), Trim. mastophtalmus ( Reinh .
Richter), Trim . polonicus n . sp., Dianops? trifolius n . sp.
IlCJIpHJI C.JIOHlVI!H
cP3J.VI€lHI BbupaoKelH 3 A€C b r a iose rB BliiAe IM ep re JIHCTbI X CJlaHI~eB C Il IPO-
1i3,OOCTlHcfI]KOS.
granulatus
(MUnster) ,
P h acop i d ae
D i an op s
IJ1Ip€I]:lCT a BJIe HibI
typhlops
CJIev:G'IO~ltMH
(Giirich) . D uct i n a
BHAaMH :
d uctifro ns
P h acops:
( R.
&
E.
Richter).
I'ep6hI -
B
,go pore lBe,gY lIJ,e H IB lJ a pHoB o6HaJ pym HBa lO T.c-H q epHible
U3IBeC'i'Hmm ,
B iKOTOiPbIX scnpesaerca Dianops typhlops (Glirich) .
Jlaroa -
06HJaiRelHlHH lP aJM€lHCl~Oro s p yca IIIOHi!lJ1.J1I<ll'OH IHa
CTe'IJKa , IHa CKJlOHHX il\:omliHbI Ip elJ:K H JIaJI'OBHl\bI.
3 1'0
oesep ncor 'KoH ~e xre -
OHJlblHIO 6HTy~1I'UIJlH03iH1ble OJIaJH~eoo­
H3lI3lOCTIHilliOBbI € o c anron. B WMHIO-<c e;pbI'X K,11!1'M eHHeBbl X H3Bec 11HHiK ax IHJaxOAHT'CH l1I00nAa
6 0 JIbffi%e OHY'I1.JleHHH naJH~bllpe H Trimeroc ephalus caecu s (G lir i c h )
usre
H IIDeMlHOI'OilIIHCTIe H -
Trim. mastophtalmus (Reinh. R ichter) . ,H 3 l~ pynH x Phacopi dae ITIlPlfBrO~HJl H C b
OTClOM Dianops typhlops (Glirich) .
P ooBHTH e
epaM e Ha IB
JIa I10 Be
HeOKOJIbKO rpa3lHHTCH
OT I11Hm rqrHo ro
C JIaH~eBO- H 3 ­
B'OC1lHIHIKOBOro, T<lJK HaJK 3):\,e c b I1'pe06JI~aIOT H,3 BeICTIHHlRln, a C13lIDl\e iB Me,Hbille . cP<lJYH fI C11HqeC,K3JH accou a an aa T<lJWKe OTnnx aerca OT TIpe,l:\bIJ(Y~HX : l~poM e oq eHlb MHOI'O,q'HcJlel-JHbIX Ph a cop ida e ,HJa,XOAHTCH T 3KHt€
'i'pHJIOOHTbI iH3 c €IM.e ikllB3J Pro etid a e .
FAMENNIAN PHACOPIDAE FROM HOLY CR O S S MTS .
B
30
f aJI€tH 3~l..\e -l#WI eIll:1U1,e
CHiHJK.l1JHlHaJIIII
3To cepsre, s aero
epaIM€tH
113BeC11HIHJIro~H
iP3G3 11T 13
C H:pOOHOBaT bUM 0 11Te:HJKOM IH3BeC'11HHi~
~C'11petH~ll bl TO.lII>KO ~Ba ,BiH~a, 11Ip 11TO,M epOPMbl C ,I'.lla3,1lJMlI1:
H
Ph . we d ek i n d i wedekindi R. &
P ro atidae
E.
113
rnpo''11HHoHO.roOl I''llOcT b
3 TOMy
snecs eecs xa
!MlHOro-
IK Q110 p b le 'Q11CymcT By IOT ill ll,€II111p a.lIbHONI CHHK.ll HiHqpl1e,
'i HC.lI'eHIlIbI.
¢ aMellOKl1 e
I'anea aau e l!(.,'C!peJ('Cl1BeHIlIO
Ji
q 'HiC.lla Ph acop i d a e
Phacops gr anulat us (M ti n-
B
Richter.
oca,Il;HlH IH3 BeC T H!HiKOOOI'O mHiHa 'p a 3 B1HT b l 3 onenyiouurx IlIY'Hlwrax :
~paHO)m H J~pyc
cornacno
HOqTH
H ¢ aM€tHC'~He oca~HlH 3aure'!1ClIOT IHe -
orcyrcrayer
IHa
m HiBe'i'Clll:I1X
a M qmIlqpoBb IX IH3I!l OO'l'HHJRaX.
Bb rp<lJiOOHb I TOOPAbL\~11 1I13B ec11HH K 1lJMiI1 'OOMHO-<C€lP OH, iPOOOBa TO H, .llIOKO.llaJ(HOH
OOPlGHlH H ep a~teH BlM€lIl,a err ,Bce
'1 H1'€I.lJbll 10,
a
11
O IlII1
sene-
Y:<liP HOl\lOOIMY ( 19QJ8~ XeHJIIOIIl,€tpoB bI H I1Opli30HT 3J(ecb m CY'l'C'l1Bye,T,
IllO Ba110 H OK pOOHIH. ITo
a
<Pa1\1I111.
He60.lWllIOH MOlIJ,HO C1'I1, AO
M. ¢ aY,IDa O q€t ll b OOI1.llI:>Il00 '11 ,Be.lIIHiKO.lI€llIHO H COXJpall~HJOC'11I1 .
s ter )
147
K.lI'HiM€l HH€ Bble 'roP H30 11T,bI C ,OOHIJIlOMep H eB bIM IBI}W IO-
T<lil{;jf{ eC 'raTTeIlj~OlPep!H(~B bIM , Illp HlH aJ( .'w m a ll\'¥DM
yome H m HlH€lMy IKap 60 1l1)'.
<lia YlIIa O 'i€tHl b ooraTa H TDpel)~p;aC:HO H 'OOXlP 1lJHlI IO CTlf. O HI3 COJ(e pmI1'T ,ro~a(mIT bI , IK.lJII1IM€ -
III~m,
oprouepacsr,
6 pa.XI110110.L\bl , I1.ll acTHiH'I3TomaOOpllbI€ , rnp l1.ll0 0 l1T bl
H3 Ph a c opid a e OOH3Jp y meH
111
OOTaTJi I1 p bl6 .
P h acops granulatus (Mu nster). B .peaynsr arre BblK.llHHII1-
H<'IHI1H H€lKOTOpbIX C,'1003 B OTJ(eJI bH bIX oOHameHiI1lflX iMO!lIlllOCT b ¢ <lL\1e11i l3 B
«o.nednerca
IB
KOB aJIH B
IHip acHlbI H
eppalla.
ITo
rp ae auax or 3 AO 7
¢aMeH
lI,BeT,
BbJpa}l{0H
anecs
Ille01~HOIPOlUJHO ii
(11957)
Y:3JP HOI..\IKO.vry
Fane aaauax
Cl'I.
,T 3J\ )J{e
,113 BeCllHflJK aMll1,
C11P'YKryp b l,
,B 'I-lie M
IK0110p b le
1.J 3iC110
aaneraior
o~p.ameHIH br.;\1Jl I
H1lJ
[]]P €JliCT<:lIB.lI<€oHbI 'BC'e IKJI I1oM e Hl1e,Bbl e
113 ''iII1C.ll a Ph a c opid a e 6bl.mH ecnpesea u :
!I13 B€lC11HIJIH aX
rqPI13 0 JlTbJ.
Dian op s typh lops (Giiric h) !11 Pro we dekindi
u.'ed ek i n d i R. & E. Richter . Bsnne 33.lI<€,raIOT C.ll<liHl..\b l I HIHvl~H e[1() IK<lpOOHa .
1I6J1oHHa OKO.lIO
Bopxoaa -
I[ OCJl€ s ropoa U\lIHp<OBOH IOOiiJHbI.
.n.OB3JHI1H, ,00 HX p e 3Y JIbTa 'l'bI
eppaHICb.
8'110
C!)3JM€l1l s necs 6bl.ll OOH3JpymelI ISH e p Bb le
B ,194 9
'
II-lie
Y:3JPOOll,KI1o:>I
II'. ClHJ Irnpol1 3.BO~I1.lI :tjlec b II'€OJIO r'l1'Ie a K l1e !I10 CJl e-
orry6JIJllJKOB3JHbI.
¢ aM0 H
aaneraer
IH a
H3BeCl1HlL ltaX
,T aK me, ·K aK H iB MBaM, OO,!DlffiJR OOpl1H ¢ aMeJia, 'OO<m H<JcIOlf..\aHCfI x e a n o-
,K.lIHM e~eBbl e ,r op l130HTbI . O Ha
1(0POBb llMlH C.lIOlIMI1 11 ' CO.L\ e pma'lIl,a lJl IllOB l1J(lJjMOMj'
ace
I[P€l~CT<1B.lI€lIIJaJ ilf3Be'Cl1IlJllKaj',1J11 iMO!lIllICtC'rl1 aKO'J l0
M, HlpaOlllblGWI1 IH ce;p b nMIH, a :3 ropn -
:10HT<lLX
n pHlflilioCb liBaeM blX
CP<lYlIlIa o seu s
OOI1.ll blHa .
30
Y:rupH01ll,HlI-DM :OO)W IOMe p l1e B b hM
113
ql1C.lla
P hacopidae
naftAell
C.llOJllM
-
TOJI bl{O
reeTJIO -3eJIeHbIM H.
Phacops gran u l at u s
(Muns ter) . fLpe.L\C 'l'aBl1TeJIli 31'01'0 iBHiJ(3J, BCTlpeqalO !lIIl1€C H IB lUllDl~HII1X IKJ]HiM€l HlH 63 bIX ,1'0 PII3C1HTax, Bhllpavl(eH b l OC'OOHMH xapallcrep l13YIO!lIII1MlI1'OH M aJIeHI:>J~lJjMI1, Illl€iCIKOJI b RO -'MlH,l "l ll1-
:\le'I1poB b~M I1 iKeepa..'10 H<lt.\1iI1.
T'OT-me
IIillJ( H3
CB e T.lID-3e.ll€lHIbIX IBOKJI IOMe p l10 Bb l X
IIlHiKOS \lJjMOOf IKeclJaurO Hb l , ,l\OCT lJjI 'aIOlIJ,l1e p 1lJ3M€jpa
18
H 3 <B€CT-
'M M . B aA ep3M€lHICilulu'Vm OC3\D1K 3 M H
.l1€i lillT C.lIaPdl,bl 1 ~)J{He [1() K 3JpOOHa .
qpe~11
OHHicaHHlbIX
uyancrax coaepcnarcs
1lJBroPOM
epa:.\1 €1HOHlI1X
Ph acopidae
l1pl1 ,BiH)¥'l !I13IJeCTIIlIbI€ ~e
panu.iu e
3
H3
Phacops granulatus (Munster) .
Tr imerocephalus caecus (G u r ich)
Dian ops ty p h l op s (Giir-ich),
n Tflil\il(€ BlJjJ(h l J(O OI1X rtop IlI e c6l1<lip y m€lH lIb l € IB
Ilonsnre :
a s nue
1l €lp€tjjHC-'10 HIH bIX
CB€IlTOHIpmuCIlllix
fop:
HALSZKA OSMOLSKA
148
Ph ac op s wedekindi w edekindi R. & E . Richter
Tl'i mer ocepha lus mas tophtal mus (Rei nh . R ich ter)
D uc ti n a d uc t i fr on s (R. & E. Rich ter).
Kpo:\le TOm YCT8IHOBJIell 1,1 'A Ba
1I0Bb lX H H)J.a,;
T ri m eroc ep h a l us polonicus n. sp.
D i an o p s? trifoli us n. sp.
HUHi e In jJH OO)J.c Hb l :.\Ha nHo :Jbl ;H OBbI X B H)J.OB .
Trim erocephal us p ol onic us n. sp .
(q m r .
5 ; I IJI. I I I, lj>li l'. 1, 2 a- c)
H olytypus: 1. G . 169.I1 .12, IKe lj>a JlOH , cp"ar . 5 H
ncr . I I !
2 a-c .
<1m I' .
St r a t um ty p i cum : CP aIMoElll , oKJI-HrMoElHtUeBb l€ CJI0 H.
Loc us t y p icus:
:K3J)J: 3€JWH~I ,
Cs enr onpxcac mre I'opu , D OJl b Il W .
onHcall Hib l O H 3 Ilom.nm.
D eriva t i o no m i nis : po l on i cus Anarno 3 . -
ocnpeeesas :
H QIlTY,p 'Ke cpaJ IOI Kl
yiHC nO: IYU\ip :YI'a;
cnepeznr
rJlBoe:lJI <lJ
O;W b HO
sa-
yrcct H X O'l€lHb
H3 MeHI'I HH blii (51'- 80"); ,H a n o.1YMecH 'I'IImllIe'P'e~IIe:\1 JIlIM 6 y c c npononroaaroe y r n y 6 JleHlHe; JIHI \CBOH
W OIB rtepec exaer Il~ eKY Up H ee rtepenae or yrn e: opnacrenr auan
'()IIHiHIHU, IC
6 0 P OOAbI .1C I1KO
BbI I' IIYTbIc , BonHYT ble ,H<lJP Y:lIiY ,
Rc¢ <WIO Ha B B~AC 6y rop K Ql!3 ,L\O BO.l b IlO 6o.1blllH..'C IB COOTHIOIIlI€lH IiIi C ')>a3 Me pa.Wf
JIOIl<lJ, p eJl;Ko !pa 3,:\l e ~ eHI Dblx ; 11 0 OOK3 :\1
H n H nlf~Hii
li e
n poveacyrosuoco
xerpa-
KO.%I\a ABa 6yropKa. T OpaJKC
H3B€Cl'Hbl.
Dianops? tr ifolius n , sp.
6;
(llm I'.
IV!.
IV,
cpHI' .
2, 3)
H ol ot y p us: 1. G . 169.1 1.16" a-(e<jJaJlO H, <1mI' . 6 H IIJ I . I V,
elmI'.
2, 3.
St ratum typ icum : q,aM€>H , xeanouepoesre MOH.
Loc us typic us:
K ClIA3 e JIbIlH, CBeHiTOWpHiI{()l'HC
D er i v a t i o n ominis: trifolius -
AnarH0 3 .
.TIHCT
-
KoHTYIP
,J{e<jJa:m u a
neneae pa: rnaoenna ClIeper,il,u
nerxo o pe3 a'IUHb l,e ;
, 6000B b [ e 'yor:I b [
I'o psr, Ilonsur a.
'IlCllllo :\u ullaIOIIl,H i:i
mlJpymHocT b lO
OHJlO~eHlIJaH, A13~bI
unupe
1'peXJlOllacT Hblii
ODOO BalHiHrfl, ee
yr.ne JI;Ba )j{J\b l unspe
HHJy11pb. To pae cx ITH!n HWin
'1eM ,y
,HiIirM 6 y ,c ODP H 3ai/llHc:\1 ~E:lfOOM
<reM Imp H n epe,l1JH€,M, 'n p H Y CTbH X CJIIHIHIIlIbl'X 6oP03 A OOI1HY T h l H
He H 3 BeC 1'll bI.
xneeepa (Trifoliumj .
H e epa JIOHI H:\l eCT EHA Jm CTa
PL AT E S
EXPLA N ATI O NS OF PLATES
PI. I
P h a cop s g ranulat u s (M Un ste r )
F i g. 1. Ceph a lon , d o rsa l v iew (I. G . 169. II . 1); X 5.
P h aco ps w ed eki n di w edeki ndi R . & E. R ich ter
Fig. 2. a Fragmen t of ce p h a l on, dor sa l v iew ; b same sp ecimen , lateral v ie w (I. G.
169. II. 6); >< 4.
T r imer ocep h a l u s mas t ophtalmus (Re in h . R ich ter)
Fig. 3. a Cephalon , d orsa l v iew; b same s p eci m e n, ante rior v ie w; c same spe cimen,
lateral v iew (I . G. 169. II. 7); X 3.1.
PI. II
Trim erocep ha lu s caecus (G Ur ich )
Fig. 1. a Enrolled specimen, d orsal vi ew ; b same specimen , a n te r ior view; c sa m e
speci men , lateral view O. G . 169. II. 8) ; X 4.7.
Fig. 2. a Internal m ould of wh ol e spe cimen , d orsal v ie w ; b same sp eci m e n, pygidium (I. G. 169 . II. »n , X 2.7.
Fig. 3. Thin section of limeston e from Lagow showin g ex oskele to ns of Trimeroce phalus cc ecus: Y 3.
P I. III
T rimer oceph altl s p olonicus n. s p .
F i g. 1. P ara t y p e, ce p h a lon flattened, d orsal view (I. G. 169. II. 26) ; X 6.4.
F ig. 2. a H olo ty p e , ce ph alo n , d or sa l vi ew ; b same specimen, anterior vi ew; c s am e
specimen , la tera l view (l G. 169. II. 12); X 8.7.
D u cti na due t if rolls R . & E. Ri ch te r
Fig. 3. Fragment of ce p ha lon , d or sal vi ew (I. G. 169. II. 20) ; X 9.5.
Pha cop s sp.
Fig . 4. Pygidium , d orsal view (1. G . 169. I I. 19); X 4.8.
P I. IV
Dian op s typ h lops (Gii r ich )
Fig. 1. Cepha lon, d orsa l v ie w (I. G . 169. I I. 14) ; X 4.7.
Fig. 4. P y gid iu m , d orsal v iew (1. G. 169. II. 15); X 8.7.
D iu n o ps ? tr ifol iu s n . sp .
Fi g. 2. H ol o type , ce p halon , d orsa l v iew (I. G. 169. II. 1,6); X 8.4.
Fig. 3. P a r a t yp e, ce p h a l o n, dorsal v ie w (I. G. 169. II. 17); Y 5.6.
ACTA P ALAEONTOLOGICA POLONICA, VOL. III
H . OSM6LSK A, P L , I
ACTA PALAEONTOLOGICA POLONICA. VOL. III
H . OSM6LSKA. PL .
rr
.ACTA PALAEONTOLOGICA POLONICA, VOL , III
H . OSMOLSKA , PL
III
ACTA PALAEONTOLOGICA POLONICA . V O L . III
1: . O Sl\~6LS K A . PL . IV